Publications

Google Scholar page

  1. Sudweeks, J., Hollingsworth, B., Blondel, D.V., Campbell, K.J., Dhole, S., Eisemann, J.D., Edwards, O., Godwin, J., Howald, G.R., Oh, K., Piaggio, A.J., Prowse, T.A.A., Ross, J.V., Saah, J.R., Shiels, A.B., Thomas, P.Q., Threadgill, D.W., Vella, M.R., Gould, F. & Lloyd, A.L. (2019) Locally Fixed Alleles: A method to localize gene drive to island populations. Sci. Rep. 9, 15821.
  2. Dhole, S., Lloyd, A.L. & Gould, F. (2019) Tethered homing gene drives: A new design for spatially restricted population replacement and suppression. Evol. Applic. 12, 1688-1702.
  3. Gould, F., Dhole, S. & Lloyd, A.L. (2019) Pest management by genetic addiction.
    Proc. Natl. Acad. Sci. 116, 5849-5851.
  4. Arnold, A. & Lloyd, A.L. (2018) An approach to periodic, time-varying parameter estimation using nonlinear filtering. Inverse Problems. 34(10), 105005. doi:10.1088/1361-6420/aad3e0
  5. ten Bosch, Q. A., Clapham, H.E., Lambrechts, L., Duong, V., Buchy, P., Althouse, B.M., Lloyd, A.L., Waller, L.A., Morrison, A.C., Kitron, U., Vazquez-Prokopec, G.M., Scott, T.W. & Perkins, T.A. (2018) Contributions from the silent majority dominate dengue virus transmission. PLOS Pathogens 14(5): e1006965. doi:10.1371/journal.ppat.1006965
  6. Gunning, C.E., Okamoto, K., Astete, H., Vasquez, G.M., Erhardt, E., Del Aguila, C., Pinedo, R., Cardenas, R., Pacheco, C., Chalco, E., Rodriguez-Ferruci, H., Scott, T.W., Lloyd, A.L., Gould, F. & Morrison, A.C. (2018) Efficacy of Aedes aegypti control by indoor Ultra Low Volume (ULV) insecticide spraying in Iquitos, Peru. PLOS Negl. Trop. Dis. 12(4): e0006378. doi:10.1371/journal.pntd.0006378
  7. Dhole, S., Vella, M.R., Lloyd, A.L. & Gould, F. (2018) Invasion and migration of spatially self-limiting gene drives: a comparative analysis. Evol. Applic. 11(5): 794-808. bioRxiv version doi:10.1111/eva.12583
  8. Yakob, L., Lloyd, A.L., Kao, R., Ferguson, H.M., Brock, P., Drakeley, C. & Bonsall, M. (2018) Plasmodium knowlesi invasion following spread by infected mosquitoes, macaques and humans. Parasitology. 145(1):101-110. doi: 10.1017/S0031182016002456
  9. Vella, M.R., Gunning, C.E., Lloyd, A.L. & Gould, F. (2017) Evaluating strategies for reversing CRISPR-Cas9 gene drives. Scientific Reports. 7: 11038. doi:10.1038/s41598-017-10633-2 PDF
  10. Hamilton, F.W., Lloyd, A.L. & Flores, K.B. (2017) Hybrid modeling and prediction of dynamical systems. PLoS Comput. Biol. 13(7): e1005655. doi:10.1371/journal.pcbi.1005655
  11. Hamilton, F., Setzer, B., Chavez, S., Tran, H. & Lloyd, A.L. (2017) Adaptive filtering for hidden node detection and tracking in networks. Chaos. 27, 073106. doi:10.1063/1.4990985
  12. Usaini, S., Lloyd, A.L., Anguelov, R. & Garba, S.M. (2017) Dynamical behavior of an epidemiological model with a demographic Allee effect. Math. Comput. Simul. 133, 311. doi:10.1016/j.matcom.2016.04.010 PDF
  13. Legros, M., Otero, M., Aznar, V.R., Solari, H., Gould, F. & Lloyd, A.L. (2016) Comparison of two detailed models of Aedes aegypti population dynamics. Ecosphere. 7(10):e01515 PDF Supp. Info. doi:10.1002/ecs2.1515
  14. Monaghan, A.J., Morin, C.W., Steinhoff, D.F., Wilhelmi, O.V., Hayden, M., Quattrochi, D.A., Reiskind, M.H., Lloyd, A.L., Smith, K., Schmidt, C.A., Scalf, P.E. & Ernst, K.C. (2016) On the seasonal occurrence and abundance of the Zika virus vector mosquito Aedes aegypti in the contiguous United States. PLoS Currents Outbreaks.
    doi:10.1371/currents.outbreaks.50dfc7f46798675fc63e7d7da563da76 LINK
  15. Okamoto, K.W., Gould, F. & Lloyd, A.L. (2016). Integrating transgenic vector manipulation with clinical interventions to manage vector-borne diseases. PLoS Comput. Biol. 12(3): e1004695. doi:10.1371/journal.pcbi.1004695 PDF
  16. Breen, M., Villeneuve, D.L., Ankley, G.T., Bencic, D., Breen, M.S., Watanabe, K.H., Lloyd, A.L. & Conolly, R.B. (2016). Computational model of the fathead minnow hypothalamic-pituitary-gonadal axis: Incorporating protein synthesis in improving predictability of responses to endocrine active chemicals. Comparat. Biochem. Physiol. C. 183-184:36-45.
    doi:10.1016/j.cbpc.2016.02.002 PDF
  17. Vazquez-Prokopec, G.M., Perkins, T.A., Waller, L.A., Lloyd, A.L., Reiner, R.C., Scott, T.W. & Kitron, U. (2016). Coupled heterogeneities and their impact on parasite transmission and control. Trends Parasitol.   32:356–367. doi:10.1016/j.pt.2016.01.001 PDF
  18. Heesterbeek, H., Anderson, R.M, Andreasen, V., Bansal, S., De Angelis, D., Dye, C., Eames, K.T.D., Edmunds, W.J., Frost, S.D.W., Funk, S., Hollingsworth, T.D., House, T., Isham, V., Klepac, P., Lessler, J., Lloyd-Smith, J.O., Metcalf, C.J.E., Mollison, D., Pellis, L., Pulliam, J.R.C., Roberts, M.G., Viboud, C. & The Isaac Newton Institute IDD Collaboration (2015). Modeling infectious disease dynamics in the complex landscape of global health. Science 347:aaa4339. doi:10.1126/science.aaa4339
  19. Lessler, J., Edmunds, W.J., Halloran, M.E., Hollingsworth, T.D. & Lloyd, A.L. (2015). Seven challenges for model-driven data collection in experimental and observational studies. Epidemics 10: 78-82. PDF doi:10.1016/j.epidem.2014.12.002
  20. Roberts, M., Andreasen, V., Lloyd, A. & Pellis, L. (2015). Nine challenges for deterministic epidemic models. Epidemics 10: 49-53. doi:10.1016/j.epidem.2014.09.006 PDF
  21. Hollingsworth, T.D., Pulliam, J.R.C., Funk, S., Truscott, J.E., Isham, V. & Lloyd, A.L. (2015). Seven challenges for modelling indirect transmission: vector-borne diseases, macroparasites and neglected tropical diseases. Epidemics 10: 16-20. doi:10.1016/j.epidem.2014.08.007 PDF
  22. Britton, T., House, T., Lloyd, A.L., Mollison, D., Riley, S. & Trapman, P. (2015). Eight challenges for stochastic epidemic models involving global transmission. Epidemics 10: 54-57. PDF
  23. Robert, M.A., Okamoto, K.W., Gould, F. & Lloyd, A.L. (2014). Anti-pathogen genes and the replacement of disease-vectoring mosquito populations: a model-based evaluation. Evol. Applic. 7:1238–1251. doi:10.1111/eva.12219 PDF Supp. Info.
  24. Guerra, C.A., Reiner, R.C., Perkins, T.A., Lindsay, S.W., Midega, J., Brady, O.J., Barker, C.M., Reisen, W.K., Harrington, L.C., Taken, W., Kitron, U., Lloyd, A.L., Hay, S.I., Scott, T.W. & Smith, D.L. (2014). An assembly of adult female mosquito mark-release-recapture data to inform vector-borne pathogen transmission models. Parasites and Vectors. 7:276. doi:10.1186/1756-3305-7-276 PDF
  25. Reiner, R.C., Stoddard, S.T., Forshey, B.M., King, A.A., Ellis, A.M., Lloyd, A.L., Long, K.C., Rocha, C., Vilcarromero, S., Astete, H, Bazan, I., Lenhart, A., Vazquez-Prokopec, G.M., Paz-Soldan, V.A., McCall, P.J., Kitron, U., Elder, J.P., Halsey, E.S., Morrison, A., Kochel, T.J., Scott. T.W. (2014). Time-varying, serotype-specific force of infection estimates for dengue virus using 12 years of longitudinal serological data from Iquitos, Peru. Proc. Natl. Acad. Sci. 111: E2694-E2702. doi:10.1073/pnas.1314933111 PDF PDF with Supp. Info.
  26. Okamoto, K.W., Robert, M.A., Gould, F. & Lloyd, A.L. (2014). Feasible introgression of an anti-pathogen transgene into an urban mosquito population without using gene-drive. PLoS Negl. Trop. Dis. 8(7): e2827. doi:10.1371/journal.pntd.0002827 PDF Supp. Info.
  27. Smith, D.L., Perkins, T.A., Reiner, R.C., Barker, C.M., Niu, T., Chaves, L.F., Ellis, A.M., George, D.B., Le Menach, A., Pulliam, J.R.C., Bisanzio, D., Buckee, C., Chiyaka, C., Cummings, D.A.T., Garcia, A.J., Gatton, M.L., Gething, P.W., Hartley, D.M., Johnston, G., Klein, E.Y., Michael, E., Lloyd, A.L., Piggot, D.M., Reisen, W.K., Ruktanonchai, N., Singh, B.K., Stoller, J., Tatem, A.J., Kitron, U., Godfray, H.C.J., Cohen, J.M., Hay, S.I. & Scott, T.W. (2014). Recasting the Theory of Mosquito-Borne Pathogen Transmission Dynamics and Control. Trans. R. Soc. Trop. Med. Hyg. 108, 185-197. DOI:10.1093/trstmh/tru026 PDF
  28. Yarmand, H., Ivy, J.S., Denton, B. & Lloyd, A.L. (2014). Optimal Two-Phase Vaccine Allocation to Geographically Different Regions under Uncertainty. Eur. J. Oper. Res. 233, 208-219.
  29. Okamoto, K.W., Robert, M.A., Lloyd, A.L. & Gould, F. (2013). A Reduce and Replace strategy for suppressing vector-borne diseases: Insights from a stochastic, spatial model. PLoS ONE 8(12):e81860. DOI:10.1371/journal.pone.0081860 PDF.
  30. Legros, M., Xu, C., Morrison, A., Scott, T.W., Lloyd, A.L. & Gould, F. (2013). Modeling the dynamics of a non-limited and a self-limited gene drive system in structured Aedes aegypti populations. PLoS ONE 8: e83354. DOI:10.1371/journal.pone.0083354 PDF
  31. Scholle, S.O., Ypma, R.J.F., Lloyd, A.L. & Koelle, K. (2013). Viral substitution rate variation can arise from the interplay between within-host and epidemiological dynamics. Am. Nat. 182, 494-513.
  32. Robert, M.A., Okamoto, K., Lloyd, A.L. & Gould, F. (2013). A Reduce and Replace strategy for suppressing vector-borne diseases: Insights from a deterministic model. PLoS ONE 8(9): e73233. DOI:10.1371/journal.pone.0073233 PDF Supp. Info.
  33. Walsh, R.K., Aguilar, C.L., Facchinelli, L., Valerio, L., Ramsey, J.M., Scott, T.W., Lloyd, A.L. & Gould, F. (2013). Regulation of Aedes aegypti population dynamics in field systems: Quantifying direct and delayed density dependence. Am. J. Trop. Med. Hyg. 89, 68-77. DOI:10.4269/ajtmh.12-0378
  34. Breen, M., Villeneuve, D.L., Ankley, G.T., Bencic, D.C., Breen, M.S., Watanabe, K.H., Lloyd, A.L. & Conolly, R.B. (2013). Developing Predictive Approaches to Characterize Adaptive Responses of the Reproductive Endocrine Axis to Aromatase Inhibition: II. Computational Modeling. Toxicol. Sci. 133, 234-247.
  35. Reiner, R.C., Perkins, T.A., Barker, C.M., Niu, T., Chaves, L.F., Ellis, A.M., George, D.B., Le Menach, A., Pulliam, J.R.C., Bisanzio, D., Buckee, C., Chiyaka, C., Cummings, D.A.T., Garcia, A.J., Gatton, M.L., Gething, P.W., Hartley, D.M., Johnston, G., Klein, E.Y., Michael, E., Lindsay, S.W., Lloyd, A.L., Piggot, D.M., Reisen, W.K., Ruktanonchai, N., Singh, B.K., Tatem, A.J., Kitron, U., Hay, S.I., Scott, T.W., & Smith, D.L. (2013). A systematic review of mathematical models of mosquito-borne pathogen transmission: 1970-2010. J. R. Soc. Interface. 10: 20120291. PDF
  36. Facchinelli, L., Valerio, L., Ramsey, J.M., Gould, F., Walsh, R.K., Bond, G., Robert, M.A., Lloyd, A.L., James, A.A., Alphey, L. & Scott, T.W. (2013). Field cage studies and progressive evaluation of genetically-engineered mosquitoes. PLoS Negl. Trop. Dis. 7(1): e2001. PDF
  37. Legros, M., Xu, C., Okamoto, K., Scott, T.W., Morrison, A.C., Lloyd, A.L. & Gould, F. (2012). Assessing the feasibility of controlling Aedes aegypti with transgenic methods: A model-based evaluation. PLoS ONE 7(12): e52235. PDF
  38. Robert, M.A., Legros, M., Facchinelli, L., Valerio, L., Ramsey, J.M., Scott, T.W., Gould, F. & Lloyd, A.L. (2012). Mathematical models as aids for design and interpretation of experiments: The case of transgenic mosquitoes. J. Med. Entomol. 49, 1177-1188.
    PDF Supplementary information: Appendix 1, Appendix 2
  39. Capaldi, A., Behrend, S., Berman, B., Smith, J. & Lloyd, A.L. (2012). Parameter estimation and uncertainty quantification for an epidemic model. Math. Biosci. Eng. 9, 553-576. PDF
  40. Durrett, R., Gleeson, J.P., Lloyd, A.L., Mucha, P.J., Shi, F., Sivakoff, D., Socolar, J. & Varghese, C. (2012). Graph fission in an evolving voter model. Proc. Natl. Acad. Sci. USA. 109, 3682-3687. PDF
  41. Walker, T., Johnson, P.H., Moreira, L.A., Iturbe-Ormaetxe, I., Frentiu, F.D., McMeniman, C.J., Leong, Y.S., Dong, Y., Axford, J., Kriesner, P., Lloyd, A.L, Ritchie, S.A., O’Neill, S.L. & Hoffmann, A.A. (2011). The wMel Wolbachia strain blocks dengue and invades caged Aedes aegypti populations. Nature 456, 450-453.
  42. Huang, Y., Lloyd, A.L., Legros, M. & Gould, F. (2011). Gene-drive into insect populations with age and spatial structure: A theoretical assessment. Evol. Applic. 4(3), 415-428.doi:10.1111/j.1752-4571.2010.00153.x
  43. Breen, M., Breen, M.S., Terasaki, N. Yamazaki, M., Lloyd, A.L. & Conolly, R.B. (2011). Mechanistic computational model of steroidogenesis in H295R cells: Role of oxysterols and cell proliferation to improve predictability of biochemical response to endocrine active chemicals. Toxicol. Sci. 123(1), 80-93.
  44. Legros, M., Magori, K., Morrison, A.C., Xu, C., Scott, T.W., Lloyd, A.L. & Gould, F. (2011). Evaluation of location-specific predictions by a detailed simulation model of Aedes aegypti populations. PLoS ONE 6(7), e22701. doi:10.1371/journal.pone.0022701 PDF
  45. Ward, C.M., Su, J.T., Huang, Y., Lloyd, A.L., Gould, F. & Hay, B.A. (2011). Medea selfish genetic elements as tools for altering traits of wild populations: A theoretical analysis. Evolution. 65(4), 1149-1162. doi:10.1111/j.1558-5646.2010.01186.x
  46. Xu, C., Legros, M., Gould, F. & Lloyd, A.L. (2010). Understanding uncertainties in model-based predictions of the population dynamics of Aedes aegypti. PLoS Negl. Trop. Dis. 4(9), e830. doi:10.1371/journal.pntd.0000830 PDF Supplementary information
  47. Arauz, L.F., Neufeld, K.N., Lloyd, A.L. & Ojiambo, P.S. (2010). Quantitative models for germination and infection of Psuedoperonospora cubensis in response to temperature and duration of leaf wetness. Phytopathology 100, 959-967.
  48. Conlan, A.J.K., Rohani, P., Lloyd, A.L., Keeling, M. & Grenfell, B.T. (2010). Resolving the impact of waiting time distributions on the persistence of measles. J. R. Soc. Interface. 7, 561-571.
  49. Magori, K., Legros, M., Puente, M.E., Focks, D.A., Scott, T.W., Lloyd, A.L. & Gould, F. (2009). Skeeter Buster: a stochastic, spatially-explicit modeling tool for studying Aedes aegypti population replacement and population suppression strategies. PLoS Negl. Trop. Dis. 3, e508. doi:10.1371/journal.pntd.0000508 PDF Supplementary Information.
  50. Sattenspiel, S., with contributions from A. Lloyd, (2009). The Geographic Spread of Infectious Diseases. Models and Applications. Princeton University Press.
  51. Lloyd, A.L. & Sattenspiel, L. (2009). Spatiotemporal Dynamics of Measles: Synchrony and Persistence in a Disease Metapopulation. In: Spatial Ecology, S. Cantrell, C. Cosner and S. Ruan (eds.), Chapman and Hall.
  52. Cintron-Arias, A., Banks, H.T., Capaldi, A. & Lloyd, A.L. (2009). A sensitivity matrix based methodology for inverse problem formulation. J. Inv. Ill-posed Problems. 17, 545-564.
  53. Lloyd, A.L. (2009). Sensitivity of Model-Based Epidemiological Parameter Estimation to Model Assumptions. Chapter 6, pages 123-141. In: Mathematical and Statistical Estimation Approaches in Epidemiology, G. Chowell, M. Hyman, L. Bettencourt and C. Castillo-Chavez (eds.), Springer.
  54. Legros, M., Lloyd, A.L., Huang, Y. & Gould, F. (2009). Density-dependent intraspecific competition in the larval stage of Aedes aegypti (Diptera: Culicidae): Revisiting the current paradigm. J. Med. Entomol. 46, 409-419.
  55. Huang, Y., Lloyd, A.L., Legros, M. & Gould, F. (2009). Invasion of selfish genetic elements into age-structured populations. Evol. Applications 2, 143-159. doi:10.1111/j.1752-4571.2008.00049.x PDF
  56. Cintron-Arias, A., Castillo-Chavez, C., Bettencourt, L.M.A., Lloyd, A.L. & Banks, H.T. (2009). The estimation of the effective reproductive number from disease outbreak data. Math. Biosci. Eng. 6, 261-282. PDF
  57. Gould, F., Huang, Y., Legros, M. & Lloyd, A.L. (2008). A Killer-Rescue system for self-limiting gene-drive of anti-pathogen constructs. Proc. Roy. Soc. Lond. B. 275, 2823-2829. PDF Supplementary information.
  58. Lloyd, A.L., Zhang, J. & Root, A.M. (2007). Stochasticity and heterogeneity in host-vector models. J. Roy. Soc. Interface 4, 851-863. PDF Supplementary information.
  59. Huang, Y., Magori, K., Lloyd, A.L. & Gould, F. (2007). Introducing transgenes into insect populations using combined gene-drive strategies: Modeling and analysis. Insect Biochem. Molecular Biol. 37, 1054-1063. PDF
  60. Bai, P., Banks, H.T., Dediu, S., Govan, A.Y., Last, M., Lloyd, A.L., Nguyen, H.K., Olufsen, M.S., Rempala, G. & Slenning, B.D. (2007). Stochastic and deterministic models for agricultural production networks. Math. Biosci. Eng. 4, 373-402. PDF
  61. Huang, Y., Magori, K., Lloyd, A.L. & Gould, F. (2007). Introducing desirable transgenes into insect populations using Y-linked meiotic drive- A theoretical assessment. Evolution 61, 717-726. PDF
  62. Lloyd, A.L. & Valeika, S. (2007). Network Models in Epidemiology: An Overview. In: Complex Population Dynamics: Nonlinear Modeling in Ecology, Epidemiology and Genetics, B. Blasius, J. Kurths and L. Stone (eds.), World Scientific. PDF (uncorrected preprint)
  63. Lloyd, A.L., Valeika, S. & Cintron-Arias, A. (2006) Infection Dynamics on Small-World Networks. In: Modeling the Dynamics of Human Diseases: Emerging Paradigms and Challenges, A.B. Gumel, C. Castillo-Chavez and D.P. Clemence (eds.), AMS. PDF (uncorrected preprint)
  64. Lloyd, A.L. & Wodarz, D. (2006). Drug Resistance in Acute Viral Infections: Rhinovirus as a Case Study. In: Disease evolution: Models, Concepts, and Data Analyses, Z. Feng, U. Diekmann and S. Levin (eds.), AMS/DIMACS. PDF (uncorrected preprint)
  65. Sabouri, A.H., Saito, M., Lloyd, A.L., Vine, A.M., Witkover, A.W., Furukawa, Y., Izumo, S., Marshall, S.E.F, Usuku, K., Bangham, C.R.M. & Osame, M. (2004) IL-10 Promoter gene single nucleotide polymorphism determines both provirus load of human T-lymphotropic virus type I (HTLV-I) in infected individuals and the risk of HTLV-I associated myelopathy/tropical spastic paraparesis (HAM/TSP). J. Inf. Dis. 190, 1279-1285. PDF
  66. Wodarz, D., & Lloyd, A.L. (2004). Immune responses and the emergence of drug-resistant virus strains in vivo. Proc. Roy. Soc. Lond. B. 271, 1101-1109. PDF
  67. Lloyd, A.L. (2004). Estimating variability in models for recurrent epidemics: assessing the use of moment closure techniques. Theor. Popul. Biol. 65, 49-65. PDF
  68. Lloyd, A.L., & Jansen, V.A.A. (2004). Spatiotemporal dynamics of epidemics: synchrony in metapopulation models. Math. Biosci. 188, 1-16. PDF
  69. Vine, A.M., Witkover, A.D., Lloyd, A.L., Jeffery, K.J., Siddiqui, A., Marshall, S.E., Bunce, M., Eiraku, N., Izumo, S., Usuku, K., Osame, M., & Bangham, C.R. (2002). Polygenic control of Human T Lymphotropic Virus type I (HTLV-I) provirus load and the risk of HTLV-I-associated Myelopathy/Tropical Spastic Paraparesis. J. Infect. Dis. 186, 932-9. PDF
  70. Gravenor, M.B., Lloyd, A.L., Kremsner, P.G., Missinou, M.A., English, M., Marsh, K., & Kwiatkowski, D. (2002). A model for estimating total parasite load in falciparum malaria patients. J. Theor. Biol. 217 , 137-148. PDF
  71. May, R.M., & Lloyd, A.L. (2001). Infection dynamics on scale-free networks. Phys. Rev. E. 64, 066112. PDF
  72. Lloyd, A.L. (2001). Realistic distributions of infectious periods in epidemic models: changing patterns of persistence and dynamics. Theor. Popul. Biol. 60, 59-71. PDF
  73. Lloyd, A.L., & May, R.M. (2001). How viruses spread among computers and people. Science 292, 1316-1317.
  74. Lloyd, A.L. (2001). Destabilization of epidemic models with the inclusion of realistic distributions of infectious periods. Proc. Roy. Soc. Lond. B. 268, 985-993. PDF online appendix
  75. Lloyd, A.L. (2001). The dependence of viral parameter estimates on the assumed viral life cycle: limitations of studies of viral load data. Proc. Roy. Soc. Lond. B. 268, 847-854. PDF
  76. Jeffery, K.J.M., Siddiqui, A.A., Bunce, M., Lloyd, A.L., Vine, A.M., Witkover, A.D., Izumo, S., Usuku, K., Welsh, K.I., Osame, M., & Bangham, C.R.M. (2000). The influence of HLA class I alleles and heterozygosity on the outcome of human T cell lymphotropic virus type T infection. J. Immunol. 165, 7278-7284. PDF
  77. Jansen, V.A.A., & Lloyd, A.L. (2000). Local stability analysis of spatially homogeneous solutions of multi-patch systems. J. Math. Biol. 41, 232-252. PDF
  78. Masel, J., Arnaout, R.A., O’Brien, T.R., Goedert, J.J., & Lloyd, A.L. (2000). Fluctuations in HIV-1 viral load are correlated to CD4(+) T- lymphocyte count during the natural course of infection. JAIDS 23, 375-379.
  79. Barchet, W., Oehen, S., Klenerman, P., Wodarz, D., Bocharov, G., Lloyd, A.L., Nowak, M.A., Hengartner, H., Zinkernagel, R.M., & Ehl, S. (2000). Direct quantitation of rapid elimination of viral antigen-positive lymphocytes by antiviral CD8(+) T cells in vivo. Eur. J. Immunol. 30, 1356-1363. PDF
  80. Lloyd, A.L., & May, R.M. (1999) Synchronicity, chaos and population cycles: spatial coherence in an uncertain world. Trends Ecol. Evol. 14, 417-418. PDF
  81. Arnaout, R.A., Lloyd, A.L., O’Brien, T.R., Goedert, J.J., Leonard, J.M., & Nowak, M.A. (1999). A simple relationship between viral load and survival time in HIV-1 infection. Proc. Natl. Acad. Sci. USA 96, 11549-11553. PDF
  82. Haydon, D.T., & Lloyd, A.L. (1999). On the origins of the Lotka-Volterra Equations. Bull. Ecol. Soc. America 80, 205-206. PDF
  83. Jeffery, K.J.M., Usuku, K., Hall, S.E., Matsumoto, W., Taylor, G.P., Procter, J., Bunce, M., Ogg, G.S., Welsh, K.I., Weber, J.N., Lloyd, A.L., Nowak, M.A., Nagai, M., Kodama, D., Izumo, S., Osame, M., & Bangham, C.R.M. (1999). HLA alleles determine human T-lymphotropic virus-I (HTLV-I) proviral load and the risk of HTLV-I-associated myelopathy. Proc. Natl. Acad. Sci. USA 96, 3848-3853. PDF
  84. Wodarz, D., Lloyd, A.L., Jansen, V.A.A., & Nowak, M.A. (1999). Dynamics of macrophage and T cell infection by HIV. J. Theor. Biol. 196, 101-113. PDF
  85. Gravenor, M.B., & Lloyd, A.L. (1998). Reply to: Models for the in-host dynamics of malaria revisited: errors in some basic models lead to large overestimates of growth rates. Parasitology 117, 409-410. PDF
  86. Chun, T.W., Stuyver, L., Mizell, S.B., Ehler, L.A., Mican, J.A.M., Baseler, M., Lloyd, A.L., Nowak, M.A., & Fauci, A.S. (1997). Presence of an inducible HIV-1 latent reservoir during highly active antiretroviral therapy. Proc. Natl. Acad. Sci. USA 94, 13193-13197. PDF
  87. Lifson, J.D., Nowak, M.A., Goldstein, S., Rossio, J.L., Kinter, A., Vasquez, G., Wiltrout, T.A., Brown, C., Schneider, D., Wahl, L., Lloyd, A.L., Williams, J., Elkins, W.R., Fauci, A.S., & Hirsch, V.M. (1997). The extent of early viral replication is a critical determinant of the natural history of simian immunodeficiency virus infection. J. Virol. 71, 9508-9514. PDF
  88. Nowak, M.A., Lloyd, A.L., Vasquez, G.M., Wiltrout, T.A., Wahl, L.M., Bischofberger, N., Williams, J., Kinter, A., Fauci, A.S., Hirsch, V.M., & Lifson, J.D. (1997). Viral dynamics of primary viremia and antiretroviral therapy in simian immunodeficiency virus infection. J. Virol. 71, 7518-7525. PDF
  89. Lloyd, A.L., & May, R.M. (1996) Spatial heterogeneity in epidemic models. J. Theor. Biol. 179, 1-11. PDF
  90. Lloyd, A.L., & Lloyd, D. (1995) Chaos- its significance and detection in biology. Biol. Rhythm Res. 26, 233-252.
  91. Lloyd, A.L. (1995). Computing bouts of the prisoner’s dilemma. Sci. Am. 272 (6), 110-115.
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  99. Lloyd, D., James, C.J., Lloyd, A.L., Yarlett, N., & Yarlett, N.C. (1987). The effects of oxygen on fermentation in Tritrichomonas fetus KV1 and its variant 1MR-100 with defective hydrogenosomes. J. Gen. Microbiol. 133, 1181-1186.