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  1. Vella, M.R., Gould, F. & Lloyd, A.L. (2021) Mathematical modeling of genetic pest management through female-specific lethality: Is one locus better than two? Evolutionary Applications. Preprint available at
  2. Oh, K.P., Shiels, A.B., Shiels, L., Blondel, D.V., Campbell, K.J., Saah, J.R., LLOYD, A.L., Thomas, P.Q., Gould, F., Abdo, Z., Godwin, JH.R., Piaggio, A.J. (2020). Population genomics of invasive rodents on islands: genetic consequences of colonization and prospects for localized synthetic gene drive. Evolutionary Applications.
  3. Bradshaw, W.J., Alley, E.C., Huggins, J.H., LLOYD, A.L. & Esvelt, K.M. (2021) Bidirectional contact tracing dramatically improves COVID-19 control. Nature Communications. Preprint version available at
  4. Schaber, K.L., Perkins, T.A., LLOYD, A.L., Waller, L.A., Kitron, U., Paz-Soldan, V.A., Elder, J.P., Rothman, A.L., Civitello, D.J., Elson, W.H., Morrison, A.C., Scott, T.W., Vazquez-Prokopec, G.M. (2020) Disease-driven reduction in human mobility influences human-mosquito contacts and dengue transmission dynamics. PLOS Computational Biology.
  5. Golnar. A.J., Ruell, E., Lloyd, A.L. & Pepin, K.M. (2020). Embracing dynamic models for gene drive management. Trends in Biotechnology.
  6. Hollingsworth, B., Okamoto, K. & LLOYD, A.L. (2020). After the Honeymoon, the Divorce: Unexpected Outcomes of Disease Control Measures Against Endemic Infections. PLOS Computational Biology 16 (10), e1008292.
  7. Dhole, S., LLOYD, A.L. & Gould, F. (2020) Gene drive dynamics in natural populations: The importance of density dependence, space and sex. Annual Review of Ecology, Evolution and Systematics 51, 505-531.
  8. Wells, K., Lurgi, M., Collins, B., Lucini, B., Kao, R.R., LLOYD, A.L., Frost, S.D.W. & Gravenor, M.B. (2020). Disease control across urban-rural gradients. Journal of the Royal Society Interface 17(173)20200775.
  9. Thompson, R.N., Hollingsworth, T.D., Isham, V., Arribas-Bel, D., Ashby, B., Britton, T., Challenor, P., Chappell, L.H.K, Clapham, H., Cunniffe, N.J., Dawid, A.P., Donnelly, C.A., Eggo, R.M., Funk, S., Gilbert, N., Glendinning, P., Gog, J.R., Hart, W.S., Heesterbeek, H., House, T., Keeling, M., Kiss, I.Z., Kretzschmar, M.E., Lloyd, A.L., McBryde, E.S., McCaw, J.M., McKinley, T.J., Miller, J.C., Morris, M., O’Neill, P.D., Parag, K.V., Pearson, C.A.B., Pellis, L., Pulliam, J.R.C., Ross, J.V., Scalia Tomba, G., Silverman, B.W., Struchiner, C.J., Tildesley, M.J., Trapman, T., Webb, C.R., Mollison, D. & Restif, O. (2020). Key questions for modelling COVID-19 exit strategies. Proceedings of the Royal Society B 287(1932) 20201405.
  10. Cavany, S.M., España, G., LLOYD, A.L., Waller, L.A., Kitron, U., Astete, H., Elson, W.H., Vazquez-Prokopec, G.M., Scott, T.W., Morrison, A.C., Reiner Jr, R.C. & Perkins, T.A. (2020). Optimizing the deployment of ultra-low volume and indoor residual spraying for dengue outbreak response. PLOS Computational Biology 16(4), e1007743.
  11. Hollingsworth, B., Hawkins, P., LLOYD, A.L. & Reiskind, M.H. (2020). Efficacy and spatial extent of yard-scale control of Aedes (Stegomyia) albopictus (Diptera: Culicidae) using barrier sprays and larval habitat management. Journal of Medical Entomology 57(4), 1104-1110.
  12. LLOYD, A.L., Kitron, U., Perkins, T.A.P., Vazquez-Prokopec, G.M. & Waller, L.A. (2020). The basic reproductive number for disease systems with multiple coupled heterogeneities. Mathematical Biosciences 321, 108294.
  13. Godwin, J., Serr, M., Barnhill-Dilling, S.K., Blondel, D.V., Brown, P.R., Campbell, K., Delborne, J., LLOYD, A.L., Oh, K.P., Prowse, T.A.A. and others (2019). Rodent gene drives for conservation: opportunities and data needs. Proceedings of the Royal Society B, 286(1914), 20191606.
  14. Sudweeks, J., Hollingsworth, B., Blondel, D.V., Campbell, K.J., Dhole, S., Eisemann, J.D., Edwards, O., Godwin, J., Howald, G.R., Oh, K., Piaggio, A.J., Prowse, T.A.A., Ross, J.V., Saah, J.R., Shiels, A.B., Thomas, P.Q., Threadgill, D.W., Vella, M.R., Gould, F. & Lloyd, A.L. (2019) Locally Fixed Alleles: A method to localize gene drive to island populations. Sci. Rep. 9, 15821.
  15. Dhole, S., Lloyd, A.L. & Gould, F. (2019) Tethered homing gene drives: A new design for spatially restricted population replacement and suppression. Evol. Applic. 12, 1688-1702.
  16. Gould, F., Dhole, S. & Lloyd, A.L. (2019) Pest management by genetic addiction.
    Proc. Natl. Acad. Sci. 116, 5849-5851.
  17. Arnold, A. & Lloyd, A.L. (2018) An approach to periodic, time-varying parameter estimation using nonlinear filtering. Inverse Problems. 34(10), 105005. doi:10.1088/1361-6420/aad3e0
  18. ten Bosch, Q. A., Clapham, H.E., Lambrechts, L., Duong, V., Buchy, P., Althouse, B.M., Lloyd, A.L., Waller, L.A., Morrison, A.C., Kitron, U., Vazquez-Prokopec, G.M., Scott, T.W. & Perkins, T.A. (2018) Contributions from the silent majority dominate dengue virus transmission. PLOS Pathogens 14(5): e1006965. doi:10.1371/journal.ppat.1006965
  19. Gunning, C.E., Okamoto, K., Astete, H., Vasquez, G.M., Erhardt, E., Del Aguila, C., Pinedo, R., Cardenas, R., Pacheco, C., Chalco, E., Rodriguez-Ferruci, H., Scott, T.W., Lloyd, A.L., Gould, F. & Morrison, A.C. (2018) Efficacy of Aedes aegypti control by indoor Ultra Low Volume (ULV) insecticide spraying in Iquitos, Peru. PLOS Negl. Trop. Dis. 12(4): e0006378. doi:10.1371/journal.pntd.0006378
  20. Dhole, S., Vella, M.R., Lloyd, A.L. & Gould, F. (2018) Invasion and migration of spatially self-limiting gene drives: a comparative analysis. Evol. Applic. 11(5): 794-808. bioRxiv version doi:10.1111/eva.12583
  21. Yakob, L., Lloyd, A.L., Kao, R., Ferguson, H.M., Brock, P., Drakeley, C. & Bonsall, M. (2018) Plasmodium knowlesi invasion following spread by infected mosquitoes, macaques and humans. Parasitology. 145(1):101-110. doi: 10.1017/S0031182016002456
  22. Vella, M.R., Gunning, C.E., Lloyd, A.L. & Gould, F. (2017) Evaluating strategies for reversing CRISPR-Cas9 gene drives. Scientific Reports. 7: 11038. doi:10.1038/s41598-017-10633-2 PDF
  23. Hamilton, F.W., Lloyd, A.L. & Flores, K.B. (2017) Hybrid modeling and prediction of dynamical systems. PLoS Comput. Biol. 13(7): e1005655. doi:10.1371/journal.pcbi.1005655
  24. Hamilton, F., Setzer, B., Chavez, S., Tran, H. & Lloyd, A.L. (2017) Adaptive filtering for hidden node detection and tracking in networks. Chaos. 27, 073106. doi:10.1063/1.4990985
  25. Usaini, S., Lloyd, A.L., Anguelov, R. & Garba, S.M. (2017) Dynamical behavior of an epidemiological model with a demographic Allee effect. Math. Comput. Simul. 133, 311. doi:10.1016/j.matcom.2016.04.010 PDF
  26. Legros, M., Otero, M., Aznar, V.R., Solari, H., Gould, F. & Lloyd, A.L. (2016) Comparison of two detailed models of Aedes aegypti population dynamics. Ecosphere. 7(10):e01515 PDF Supp. Info. doi:10.1002/ecs2.1515
  27. Monaghan, A.J., Morin, C.W., Steinhoff, D.F., Wilhelmi, O.V., Hayden, M., Quattrochi, D.A., Reiskind, M.H., Lloyd, A.L., Smith, K., Schmidt, C.A., Scalf, P.E. & Ernst, K.C. (2016) On the seasonal occurrence and abundance of the Zika virus vector mosquito Aedes aegypti in the contiguous United States. PLoS Currents Outbreaks.
    doi:10.1371/currents.outbreaks.50dfc7f46798675fc63e7d7da563da76 LINK
  28. Okamoto, K.W., Gould, F. & Lloyd, A.L. (2016). Integrating transgenic vector manipulation with clinical interventions to manage vector-borne diseases. PLoS Comput. Biol. 12(3): e1004695. doi:10.1371/journal.pcbi.1004695 PDF
  29. Breen, M., Villeneuve, D.L., Ankley, G.T., Bencic, D., Breen, M.S., Watanabe, K.H., Lloyd, A.L. & Conolly, R.B. (2016). Computational model of the fathead minnow hypothalamic-pituitary-gonadal axis: Incorporating protein synthesis in improving predictability of responses to endocrine active chemicals. Comparat. Biochem. Physiol. C. 183-184:36-45.
    doi:10.1016/j.cbpc.2016.02.002 PDF
  30. Vazquez-Prokopec, G.M., Perkins, T.A., Waller, L.A., Lloyd, A.L., Reiner, R.C., Scott, T.W. & Kitron, U. (2016). Coupled heterogeneities and their impact on parasite transmission and control. Trends Parasitol.   32:356–367. doi:10.1016/ PDF
  31. Heesterbeek, H., Anderson, R.M, Andreasen, V., Bansal, S., De Angelis, D., Dye, C., Eames, K.T.D., Edmunds, W.J., Frost, S.D.W., Funk, S., Hollingsworth, T.D., House, T., Isham, V., Klepac, P., Lessler, J., Lloyd-Smith, J.O., Metcalf, C.J.E., Mollison, D., Pellis, L., Pulliam, J.R.C., Roberts, M.G., Viboud, C. & The Isaac Newton Institute IDD Collaboration (2015). Modeling infectious disease dynamics in the complex landscape of global health. Science 347:aaa4339. doi:10.1126/science.aaa4339
  32. Lessler, J., Edmunds, W.J., Halloran, M.E., Hollingsworth, T.D. & Lloyd, A.L. (2015). Seven challenges for model-driven data collection in experimental and observational studies. Epidemics 10: 78-82. PDF doi:10.1016/j.epidem.2014.12.002
  33. Roberts, M., Andreasen, V., Lloyd, A. & Pellis, L. (2015). Nine challenges for deterministic epidemic models. Epidemics 10: 49-53. doi:10.1016/j.epidem.2014.09.006 PDF
  34. Hollingsworth, T.D., Pulliam, J.R.C., Funk, S., Truscott, J.E., Isham, V. & Lloyd, A.L. (2015). Seven challenges for modelling indirect transmission: vector-borne diseases, macroparasites and neglected tropical diseases. Epidemics 10: 16-20. doi:10.1016/j.epidem.2014.08.007 PDF
  35. Britton, T., House, T., Lloyd, A.L., Mollison, D., Riley, S. & Trapman, P. (2015). Eight challenges for stochastic epidemic models involving global transmission. Epidemics 10: 54-57. PDF
  36. Robert, M.A., Okamoto, K.W., Gould, F. & Lloyd, A.L. (2014). Anti-pathogen genes and the replacement of disease-vectoring mosquito populations: a model-based evaluation. Evol. Applic. 7:1238–1251. doi:10.1111/eva.12219 PDF Supp. Info.
  37. Guerra, C.A., Reiner, R.C., Perkins, T.A., Lindsay, S.W., Midega, J., Brady, O.J., Barker, C.M., Reisen, W.K., Harrington, L.C., Taken, W., Kitron, U., Lloyd, A.L., Hay, S.I., Scott, T.W. & Smith, D.L. (2014). An assembly of adult female mosquito mark-release-recapture data to inform vector-borne pathogen transmission models. Parasites and Vectors. 7:276. doi:10.1186/1756-3305-7-276 PDF
  38. Reiner, R.C., Stoddard, S.T., Forshey, B.M., King, A.A., Ellis, A.M., Lloyd, A.L., Long, K.C., Rocha, C., Vilcarromero, S., Astete, H, Bazan, I., Lenhart, A., Vazquez-Prokopec, G.M., Paz-Soldan, V.A., McCall, P.J., Kitron, U., Elder, J.P., Halsey, E.S., Morrison, A., Kochel, T.J., Scott. T.W. (2014). Time-varying, serotype-specific force of infection estimates for dengue virus using 12 years of longitudinal serological data from Iquitos, Peru. Proc. Natl. Acad. Sci. 111: E2694-E2702. doi:10.1073/pnas.1314933111 PDF PDF with Supp. Info.
  39. Okamoto, K.W., Robert, M.A., Gould, F. & Lloyd, A.L. (2014). Feasible introgression of an anti-pathogen transgene into an urban mosquito population without using gene-drive. PLoS Negl. Trop. Dis. 8(7): e2827. doi:10.1371/journal.pntd.0002827 PDF Supp. Info.
  40. Smith, D.L., Perkins, T.A., Reiner, R.C., Barker, C.M., Niu, T., Chaves, L.F., Ellis, A.M., George, D.B., Le Menach, A., Pulliam, J.R.C., Bisanzio, D., Buckee, C., Chiyaka, C., Cummings, D.A.T., Garcia, A.J., Gatton, M.L., Gething, P.W., Hartley, D.M., Johnston, G., Klein, E.Y., Michael, E., Lloyd, A.L., Piggot, D.M., Reisen, W.K., Ruktanonchai, N., Singh, B.K., Stoller, J., Tatem, A.J., Kitron, U., Godfray, H.C.J., Cohen, J.M., Hay, S.I. & Scott, T.W. (2014). Recasting the Theory of Mosquito-Borne Pathogen Transmission Dynamics and Control. Trans. R. Soc. Trop. Med. Hyg. 108, 185-197. DOI:10.1093/trstmh/tru026 PDF
  41. Yarmand, H., Ivy, J.S., Denton, B. & Lloyd, A.L. (2014). Optimal Two-Phase Vaccine Allocation to Geographically Different Regions under Uncertainty. Eur. J. Oper. Res. 233, 208-219.
  42. Okamoto, K.W., Robert, M.A., Lloyd, A.L. & Gould, F. (2013). A Reduce and Replace strategy for suppressing vector-borne diseases: Insights from a stochastic, spatial model. PLoS ONE 8(12):e81860. DOI:10.1371/journal.pone.0081860 PDF.
  43. Legros, M., Xu, C., Morrison, A., Scott, T.W., Lloyd, A.L. & Gould, F. (2013). Modeling the dynamics of a non-limited and a self-limited gene drive system in structured Aedes aegypti populations. PLoS ONE 8: e83354. DOI:10.1371/journal.pone.0083354 PDF
  44. Scholle, S.O., Ypma, R.J.F., Lloyd, A.L. & Koelle, K. (2013). Viral substitution rate variation can arise from the interplay between within-host and epidemiological dynamics. Am. Nat. 182, 494-513.
  45. Robert, M.A., Okamoto, K., Lloyd, A.L. & Gould, F. (2013). A Reduce and Replace strategy for suppressing vector-borne diseases: Insights from a deterministic model. PLoS ONE 8(9): e73233. DOI:10.1371/journal.pone.0073233 PDF Supp. Info.
  46. Walsh, R.K., Aguilar, C.L., Facchinelli, L., Valerio, L., Ramsey, J.M., Scott, T.W., Lloyd, A.L. & Gould, F. (2013). Regulation of Aedes aegypti population dynamics in field systems: Quantifying direct and delayed density dependence. Am. J. Trop. Med. Hyg. 89, 68-77. DOI:10.4269/ajtmh.12-0378
  47. Breen, M., Villeneuve, D.L., Ankley, G.T., Bencic, D.C., Breen, M.S., Watanabe, K.H., Lloyd, A.L. & Conolly, R.B. (2013). Developing Predictive Approaches to Characterize Adaptive Responses of the Reproductive Endocrine Axis to Aromatase Inhibition: II. Computational Modeling. Toxicol. Sci. 133, 234-247.
  48. Reiner, R.C., Perkins, T.A., Barker, C.M., Niu, T., Chaves, L.F., Ellis, A.M., George, D.B., Le Menach, A., Pulliam, J.R.C., Bisanzio, D., Buckee, C., Chiyaka, C., Cummings, D.A.T., Garcia, A.J., Gatton, M.L., Gething, P.W., Hartley, D.M., Johnston, G., Klein, E.Y., Michael, E., Lindsay, S.W., Lloyd, A.L., Piggot, D.M., Reisen, W.K., Ruktanonchai, N., Singh, B.K., Tatem, A.J., Kitron, U., Hay, S.I., Scott, T.W., & Smith, D.L. (2013). A systematic review of mathematical models of mosquito-borne pathogen transmission: 1970-2010. J. R. Soc. Interface. 10: 20120291. PDF
  49. Facchinelli, L., Valerio, L., Ramsey, J.M., Gould, F., Walsh, R.K., Bond, G., Robert, M.A., Lloyd, A.L., James, A.A., Alphey, L. & Scott, T.W. (2013). Field cage studies and progressive evaluation of genetically-engineered mosquitoes. PLoS Negl. Trop. Dis. 7(1): e2001. PDF
  50. Legros, M., Xu, C., Okamoto, K., Scott, T.W., Morrison, A.C., Lloyd, A.L. & Gould, F. (2012). Assessing the feasibility of controlling Aedes aegypti with transgenic methods: A model-based evaluation. PLoS ONE 7(12): e52235. PDF
  51. Robert, M.A., Legros, M., Facchinelli, L., Valerio, L., Ramsey, J.M., Scott, T.W., Gould, F. & Lloyd, A.L. (2012). Mathematical models as aids for design and interpretation of experiments: The case of transgenic mosquitoes. J. Med. Entomol. 49, 1177-1188.
    PDF Supplementary information: Appendix 1, Appendix 2
  52. Capaldi, A., Behrend, S., Berman, B., Smith, J. & Lloyd, A.L. (2012). Parameter estimation and uncertainty quantification for an epidemic model. Math. Biosci. Eng. 9, 553-576. PDF
  53. Durrett, R., Gleeson, J.P., Lloyd, A.L., Mucha, P.J., Shi, F., Sivakoff, D., Socolar, J. & Varghese, C. (2012). Graph fission in an evolving voter model. Proc. Natl. Acad. Sci. USA. 109, 3682-3687. PDF
  54. Walker, T., Johnson, P.H., Moreira, L.A., Iturbe-Ormaetxe, I., Frentiu, F.D., McMeniman, C.J., Leong, Y.S., Dong, Y., Axford, J., Kriesner, P., Lloyd, A.L, Ritchie, S.A., O’Neill, S.L. & Hoffmann, A.A. (2011). The wMel Wolbachia strain blocks dengue and invades caged Aedes aegypti populations. Nature 456, 450-453.
  55. Huang, Y., Lloyd, A.L., Legros, M. & Gould, F. (2011). Gene-drive into insect populations with age and spatial structure: A theoretical assessment. Evol. Applic. 4(3), 415-428.doi:10.1111/j.1752-4571.2010.00153.x
  56. Breen, M., Breen, M.S., Terasaki, N. Yamazaki, M., Lloyd, A.L. & Conolly, R.B. (2011). Mechanistic computational model of steroidogenesis in H295R cells: Role of oxysterols and cell proliferation to improve predictability of biochemical response to endocrine active chemicals. Toxicol. Sci. 123(1), 80-93.
  57. Legros, M., Magori, K., Morrison, A.C., Xu, C., Scott, T.W., Lloyd, A.L. & Gould, F. (2011). Evaluation of location-specific predictions by a detailed simulation model of Aedes aegypti populations. PLoS ONE 6(7), e22701. doi:10.1371/journal.pone.0022701 PDF
  58. Ward, C.M., Su, J.T., Huang, Y., Lloyd, A.L., Gould, F. & Hay, B.A. (2011). Medea selfish genetic elements as tools for altering traits of wild populations: A theoretical analysis. Evolution. 65(4), 1149-1162. doi:10.1111/j.1558-5646.2010.01186.x
  59. Xu, C., Legros, M., Gould, F. & Lloyd, A.L. (2010). Understanding uncertainties in model-based predictions of the population dynamics of Aedes aegypti. PLoS Negl. Trop. Dis. 4(9), e830. doi:10.1371/journal.pntd.0000830 PDF Supplementary information
  60. Arauz, L.F., Neufeld, K.N., Lloyd, A.L. & Ojiambo, P.S. (2010). Quantitative models for germination and infection of Psuedoperonospora cubensis in response to temperature and duration of leaf wetness. Phytopathology 100, 959-967.
  61. Conlan, A.J.K., Rohani, P., Lloyd, A.L., Keeling, M. & Grenfell, B.T. (2010). Resolving the impact of waiting time distributions on the persistence of measles. J. R. Soc. Interface. 7, 561-571.
  62. Magori, K., Legros, M., Puente, M.E., Focks, D.A., Scott, T.W., Lloyd, A.L. & Gould, F. (2009). Skeeter Buster: a stochastic, spatially-explicit modeling tool for studying Aedes aegypti population replacement and population suppression strategies. PLoS Negl. Trop. Dis. 3, e508. doi:10.1371/journal.pntd.0000508 PDF Supplementary Information.
  63. Sattenspiel, S., with contributions from A. Lloyd, (2009). The Geographic Spread of Infectious Diseases. Models and Applications. Princeton University Press.
  64. Lloyd, A.L. & Sattenspiel, L. (2009). Spatiotemporal Dynamics of Measles: Synchrony and Persistence in a Disease Metapopulation. In: Spatial Ecology, S. Cantrell, C. Cosner and S. Ruan (eds.), Chapman and Hall.
  65. Cintron-Arias, A., Banks, H.T., Capaldi, A. & Lloyd, A.L. (2009). A sensitivity matrix based methodology for inverse problem formulation. J. Inv. Ill-posed Problems. 17, 545-564.
  66. Lloyd, A.L. (2009). Sensitivity of Model-Based Epidemiological Parameter Estimation to Model Assumptions. Chapter 6, pages 123-141. In: Mathematical and Statistical Estimation Approaches in Epidemiology, G. Chowell, M. Hyman, L. Bettencourt and C. Castillo-Chavez (eds.), Springer.
  67. Legros, M., Lloyd, A.L., Huang, Y. & Gould, F. (2009). Density-dependent intraspecific competition in the larval stage of Aedes aegypti (Diptera: Culicidae): Revisiting the current paradigm. J. Med. Entomol. 46, 409-419.
  68. Huang, Y., Lloyd, A.L., Legros, M. & Gould, F. (2009). Invasion of selfish genetic elements into age-structured populations. Evol. Applications 2, 143-159. doi:10.1111/j.1752-4571.2008.00049.x PDF
  69. Cintron-Arias, A., Castillo-Chavez, C., Bettencourt, L.M.A., Lloyd, A.L. & Banks, H.T. (2009). The estimation of the effective reproductive number from disease outbreak data. Math. Biosci. Eng. 6, 261-282. PDF
  70. Gould, F., Huang, Y., Legros, M. & Lloyd, A.L. (2008). A Killer-Rescue system for self-limiting gene-drive of anti-pathogen constructs. Proc. Roy. Soc. Lond. B. 275, 2823-2829. PDF Supplementary information.
  71. Lloyd, A.L., Zhang, J. & Root, A.M. (2007). Stochasticity and heterogeneity in host-vector models. J. Roy. Soc. Interface 4, 851-863. PDF Supplementary information.
  72. Huang, Y., Magori, K., Lloyd, A.L. & Gould, F. (2007). Introducing transgenes into insect populations using combined gene-drive strategies: Modeling and analysis. Insect Biochem. Molecular Biol. 37, 1054-1063. PDF
  73. Bai, P., Banks, H.T., Dediu, S., Govan, A.Y., Last, M., Lloyd, A.L., Nguyen, H.K., Olufsen, M.S., Rempala, G. & Slenning, B.D. (2007). Stochastic and deterministic models for agricultural production networks. Math. Biosci. Eng. 4, 373-402. PDF
  74. Huang, Y., Magori, K., Lloyd, A.L. & Gould, F. (2007). Introducing desirable transgenes into insect populations using Y-linked meiotic drive- A theoretical assessment. Evolution 61, 717-726. PDF
  75. Lloyd, A.L. & Valeika, S. (2007). Network Models in Epidemiology: An Overview. In: Complex Population Dynamics: Nonlinear Modeling in Ecology, Epidemiology and Genetics, B. Blasius, J. Kurths and L. Stone (eds.), World Scientific. PDF (uncorrected preprint)
  76. Lloyd, A.L., Valeika, S. & Cintron-Arias, A. (2006) Infection Dynamics on Small-World Networks. In: Modeling the Dynamics of Human Diseases: Emerging Paradigms and Challenges, A.B. Gumel, C. Castillo-Chavez and D.P. Clemence (eds.), AMS. PDF (uncorrected preprint)
  77. Lloyd, A.L. & Wodarz, D. (2006). Drug Resistance in Acute Viral Infections: Rhinovirus as a Case Study. In: Disease evolution: Models, Concepts, and Data Analyses, Z. Feng, U. Diekmann and S. Levin (eds.), AMS/DIMACS. PDF (uncorrected preprint)
  78. Sabouri, A.H., Saito, M., Lloyd, A.L., Vine, A.M., Witkover, A.W., Furukawa, Y., Izumo, S., Marshall, S.E.F, Usuku, K., Bangham, C.R.M. & Osame, M. (2004) IL-10 Promoter gene single nucleotide polymorphism determines both provirus load of human T-lymphotropic virus type I (HTLV-I) in infected individuals and the risk of HTLV-I associated myelopathy/tropical spastic paraparesis (HAM/TSP). J. Inf. Dis. 190, 1279-1285. PDF
  79. Wodarz, D., & Lloyd, A.L. (2004). Immune responses and the emergence of drug-resistant virus strains in vivo. Proc. Roy. Soc. Lond. B. 271, 1101-1109. PDF
  80. Lloyd, A.L. (2004). Estimating variability in models for recurrent epidemics: assessing the use of moment closure techniques. Theor. Popul. Biol. 65, 49-65. PDF
  81. Lloyd, A.L., & Jansen, V.A.A. (2004). Spatiotemporal dynamics of epidemics: synchrony in metapopulation models. Math. Biosci. 188, 1-16. PDF
  82. Vine, A.M., Witkover, A.D., Lloyd, A.L., Jeffery, K.J., Siddiqui, A., Marshall, S.E., Bunce, M., Eiraku, N., Izumo, S., Usuku, K., Osame, M., & Bangham, C.R. (2002). Polygenic control of Human T Lymphotropic Virus type I (HTLV-I) provirus load and the risk of HTLV-I-associated Myelopathy/Tropical Spastic Paraparesis. J. Infect. Dis. 186, 932-9. PDF
  83. Gravenor, M.B., Lloyd, A.L., Kremsner, P.G., Missinou, M.A., English, M., Marsh, K., & Kwiatkowski, D. (2002). A model for estimating total parasite load in falciparum malaria patients. J. Theor. Biol. 217 , 137-148. PDF
  84. May, R.M., & Lloyd, A.L. (2001). Infection dynamics on scale-free networks. Phys. Rev. E. 64, 066112. PDF
  85. Lloyd, A.L. (2001). Realistic distributions of infectious periods in epidemic models: changing patterns of persistence and dynamics. Theor. Popul. Biol. 60, 59-71. PDF
  86. Lloyd, A.L., & May, R.M. (2001). How viruses spread among computers and people. Science 292, 1316-1317.
  87. Lloyd, A.L. (2001). Destabilization of epidemic models with the inclusion of realistic distributions of infectious periods. Proc. Roy. Soc. Lond. B. 268, 985-993. PDF online appendix
  88. Lloyd, A.L. (2001). The dependence of viral parameter estimates on the assumed viral life cycle: limitations of studies of viral load data. Proc. Roy. Soc. Lond. B. 268, 847-854. PDF
  89. Jeffery, K.J.M., Siddiqui, A.A., Bunce, M., Lloyd, A.L., Vine, A.M., Witkover, A.D., Izumo, S., Usuku, K., Welsh, K.I., Osame, M., & Bangham, C.R.M. (2000). The influence of HLA class I alleles and heterozygosity on the outcome of human T cell lymphotropic virus type T infection. J. Immunol. 165, 7278-7284. PDF
  90. Jansen, V.A.A., & Lloyd, A.L. (2000). Local stability analysis of spatially homogeneous solutions of multi-patch systems. J. Math. Biol. 41, 232-252. PDF
  91. Masel, J., Arnaout, R.A., O’Brien, T.R., Goedert, J.J., & Lloyd, A.L. (2000). Fluctuations in HIV-1 viral load are correlated to CD4(+) T- lymphocyte count during the natural course of infection. JAIDS 23, 375-379.
  92. Barchet, W., Oehen, S., Klenerman, P., Wodarz, D., Bocharov, G., Lloyd, A.L., Nowak, M.A., Hengartner, H., Zinkernagel, R.M., & Ehl, S. (2000). Direct quantitation of rapid elimination of viral antigen-positive lymphocytes by antiviral CD8(+) T cells in vivo. Eur. J. Immunol. 30, 1356-1363. PDF
  93. Lloyd, A.L., & May, R.M. (1999) Synchronicity, chaos and population cycles: spatial coherence in an uncertain world. Trends Ecol. Evol. 14, 417-418. PDF
  94. Arnaout, R.A., Lloyd, A.L., O’Brien, T.R., Goedert, J.J., Leonard, J.M., & Nowak, M.A. (1999). A simple relationship between viral load and survival time in HIV-1 infection. Proc. Natl. Acad. Sci. USA 96, 11549-11553. PDF
  95. Haydon, D.T., & Lloyd, A.L. (1999). On the origins of the Lotka-Volterra Equations. Bull. Ecol. Soc. America 80, 205-206. PDF
  96. Jeffery, K.J.M., Usuku, K., Hall, S.E., Matsumoto, W., Taylor, G.P., Procter, J., Bunce, M., Ogg, G.S., Welsh, K.I., Weber, J.N., Lloyd, A.L., Nowak, M.A., Nagai, M., Kodama, D., Izumo, S., Osame, M., & Bangham, C.R.M. (1999). HLA alleles determine human T-lymphotropic virus-I (HTLV-I) proviral load and the risk of HTLV-I-associated myelopathy. Proc. Natl. Acad. Sci. USA 96, 3848-3853. PDF
  97. Wodarz, D., Lloyd, A.L., Jansen, V.A.A., & Nowak, M.A. (1999). Dynamics of macrophage and T cell infection by HIV. J. Theor. Biol. 196, 101-113. PDF
  98. Gravenor, M.B., & Lloyd, A.L. (1998). Reply to: Models for the in-host dynamics of malaria revisited: errors in some basic models lead to large overestimates of growth rates. Parasitology 117, 409-410. PDF
  99. Chun, T.W., Stuyver, L., Mizell, S.B., Ehler, L.A., Mican, J.A.M., Baseler, M., Lloyd, A.L., Nowak, M.A., & Fauci, A.S. (1997). Presence of an inducible HIV-1 latent reservoir during highly active antiretroviral therapy. Proc. Natl. Acad. Sci. USA 94, 13193-13197. PDF
  100. Lifson, J.D., Nowak, M.A., Goldstein, S., Rossio, J.L., Kinter, A., Vasquez, G., Wiltrout, T.A., Brown, C., Schneider, D., Wahl, L., Lloyd, A.L., Williams, J., Elkins, W.R., Fauci, A.S., & Hirsch, V.M. (1997). The extent of early viral replication is a critical determinant of the natural history of simian immunodeficiency virus infection. J. Virol. 71, 9508-9514. PDF
  101. Nowak, M.A., Lloyd, A.L., Vasquez, G.M., Wiltrout, T.A., Wahl, L.M., Bischofberger, N., Williams, J., Kinter, A., Fauci, A.S., Hirsch, V.M., & Lifson, J.D. (1997). Viral dynamics of primary viremia and antiretroviral therapy in simian immunodeficiency virus infection. J. Virol. 71, 7518-7525. PDF
  102. Lloyd, A.L., & May, R.M. (1996) Spatial heterogeneity in epidemic models. J. Theor. Biol. 179, 1-11. PDF
  103. Lloyd, A.L., & Lloyd, D. (1995) Chaos- its significance and detection in biology. Biol. Rhythm Res. 26, 233-252.
  104. Lloyd, A.L. (1995). Computing bouts of the prisoner’s dilemma. Sci. Am. 272 (6), 110-115.
  105. Lloyd, A.L. (1995). The coupled logistic map- A simple model for the effects of spatial heterogeneity on population dynamics. J. Theor. Biol. 173, 217-230. PDF
  106. Lloyd, D., & Lloyd, A.L. (1994). Hypothesis- A controlled chaotic attractor constitutes the central oscillator of the circadian clock. Biochem. Soc. Trans. 22, S322.
  107. Lloyd, A.L. (1994). Chaos and forecasting. Trends Ecol. Evol. 9, 244-245.
  108. Lloyd, A.L., & Gravenor, M.B. (1994). Linear filters and nonlinear forecasting. Proc. Roy. Soc. Lond. B 256, 157-161. PDF
  109. Mason, D.J., Lloyd, A.L. & Lloyd, D. (1993). An inexpensive infrared growth sensor array for detection of bacterial antibiotic susceptibility. FEMS Microbiol. Lett. 111, 251-254.
  110. Lloyd, A.L., & Lloyd, D. (1993). Hypothesis- the central oscillator of the circadian clock is a controlled chaotic attractor. Biosystems 29, 77-85.
  111. Lloyd, D., Lloyd, A.L., & Olsen, L.F. (1992) The cell-division cycle-a physiologically plausible dynamic model can exhibit chaotic solutions. Biosystems 27, 17-24. PDF
  112. Lloyd, D., James, C.J., Lloyd, A.L., Yarlett, N., & Yarlett, N.C. (1987). The effects of oxygen on fermentation in Tritrichomonas fetus KV1 and its variant 1MR-100 with defective hydrogenosomes. J. Gen. Microbiol. 133, 1181-1186.